Staff profile

• 2019: Research Associate; National Museum of Scotland:
• 2019: Guest Research Professor, Chinese Academy of Sciences, Nanjing:
• 2017: Distinguished Visiting Professor, China University of Geosciences, Wuhan:
• 2016: Leverhulme Fellow:
• 2016: Chair, International Commission on Stratigraphy:
• 2014: President, Palaeontological Association:
• 2014: Visiting Professor, CNRS - Université de Lille 1, France:
• 2014: Guest Professor, University of Lund:
• 2014: Elected Foreign Member of the Royal Swedish Physiographic Society:
• 2014: Member of Scientific Advisory Board, Naturalis (State Museum of Natural History), The Netherlands:
• 2014: William King Medal, National University of Ireland, Galway:
• 2013: Honorary Research Fellow, The Natural History Museum:
• 2008: Einstein Professor, Chinese Academy of Sciences:
• 2004: Elected Foreign Member of the Royal Danish Academy of Sciences and Letters:
• 2003: Crown Prince Frederik's Fund, Denmark:
• 1996: Lyell Fund from the Geological Society:
• 1986: Clough Memorial Award from the Edinburgh Geological Society:

Authored book

• Liljeroth, M., Harper, D., Carlisle, H., & Nielsen, A. Ordovician rhynchonelliformean brachiopods from Co. Waterford, SE Ireland: Palaeobiogeography of the Leinster Terrane. John Wiley and Sons
• Benton, M., & Harper, D. (2020). Introduction to Paleobiology and the Fossil Record, 2nd edition. Wiley
• Jiayu, R., Bing, H., Ren-bin, Z., & Harper, D. (2013). Latest Ordovician and earliest Silurian brachiopods succeeding the Hirnantia fauna in south-east China. Palaeontological Association. https://doi.org/10.1111/pala.12056
• Rasmussen, C., Harper, D., & Blodgett, R. (2012). Late Ordovician Brachiopods from West-Central Alaska: systematics, ecology and palaeobiogeography. Wiley
• Benton, M., & Harper, D. (2009). Introduction to Paleobiology and the Fossil Record. John Wiley and Sons

Chapter in book

• Harper, D. A. (2021). Brachiopoda. In D. Alderton, & S. A. Elias (Eds.), Encyclopedia of Geology (273-282). (2nd). Elsevier. https://doi.org/10.1016/b978-0-08-102908-4.00108-9
• Harper, D. A. (2021). Late Ordovician Extinctions. In D. Alderton, & S. A. Elias (Eds.), Encyclopedia of Geology (617-626). (2nd). Elsevier. https://doi.org/10.1016/b978-0-12-409548-9.12530-8
• Harper, D. A. (2021). Paleontology. In J. Vonk, & T. Shackelford (Eds.), Encyclopedia of Animal Cognition and Behavior. Springer Verlag. https://doi.org/10.1007/978-3-319-47829-6_527-1
• Armstrong, H., & Harper, D. (2014). An earth system approach to understanding the end-Ordovician (Hirnantian) mass extinction. In G. Keller, & A. Kerr (Eds.), Volcanism, impacts, and mass extinctions : causes and effects (287-300). Geological Society of America. https://doi.org/10.1130/2014.2505%2814%29
• Platform: North Atlantic Region. In J. Derby, R. Fritz, S. Longacre, W. Morgan, & C. Sternbach (Eds.), The great American carbonate bank: The geology and economic resources of the Cambrian-Ordovician Sauk megasequence of Laurentia (597-626). AAPG Memoir

Edited book

• Harper, D., & Servais, T. (Eds.). (2013). Early Palaeozoic Biogeography and Palaeogeography. Geological Society

Journal Article

• Murray, J., Nasheuer, H., Seoighe, C., McCormick, G., Williams, D., & Harper, D. (in press). The contribution of William King to the early development of palaeoanthropology. Irish Journal of Earth Sciences, 33, 1-16. https://doi.org/10.3318/ijes.2015.33.1
• Guo, Z., Chen, Z., & Harper, D. A. (in press). Phylogenetic and ecomorphologic diversifications of spiriferinid brachiopods after the end-Permian extinction. Paleobiology, 1-16. https://doi.org/10.1017/pab.2020.34
• Penn-Clarke, C. R., & Harper, D. A. (in press). Early−Middle Devonian brachiopod provincialism and bioregionalization at high latitudes: A case study from southwestern Gondwana. GSA Bulletin, https://doi.org/10.1130/b35670.1
• Harper, D., Zhan, R., & Jin, J. (in press). The Great Ordovician Biodiversification Event: Reviewing two decades of research on diversity’s big bang illustrated by mainly brachiopod data. Palaeoworld, 24(1-2), 75-85. https://doi.org/10.1016/j.palwor.2015.03.003
• Wu, S., Reitner, J., Harper, D. A. T., Yu, J., & Chen, Z. (2024). New keratose sponges after the end-Permian extinction provide insights into biotic recoveries. Geobiology, 22(1), Article e12582. https://doi.org/10.1111/gbi.12582
• Harper, D. A. T. (2024). Late Ordovician Mass Extinction: Earth, fire and ice. National Science Review, 11(1), Article nwad319. https://doi.org/10.1093/nsr/nwad319
• Park, T. S., Nielsen, M. L., Parry, L. A., Sørensen, M. V., Lee, M., Kihm, J., …Vinther, J. (2024). A giant stem-group chaetognath. Science Advances, 10(1), Article eadi6678. https://doi.org/10.1126/sciadv.adi6678
• Jin, J., Rasmussen, C. M. Ø., Sheehan, P. M., & Harper, D. A. T. (2024). Late Ordovician and early Silurian virgianid and stricklandioid brachiopods from North Greenland: implications for a warm‐water faunal province. Papers in Palaeontology, 10(1), Article e1544. https://doi.org/10.1002/spp2.1544
• Harper, D. A., & Bates, D. E. (2023). Middle Ordovician brachiopods from Tagoat, Co. Wexford, SE Ireland: Dapingian diversity drivers. Geobios, 81, 85-100. https://doi.org/10.1016/j.geobios.2023.06.006
• Penn-Clarke, C. R., & Harper, D. A. (2023). The rise and fall of the Malvinoxhosan (Malvinokaffric) bioregion in South Africa: Evidence for Early-Middle Devonian biocrises at the South Pole. Earth-Science Reviews, 246, Article 104595. https://doi.org/10.1016/j.earscirev.2023.104595
• Servais, T., Cascales-Miñana, B., Harper, D. A., Lefebvre, B., Munnecke, A., Wang, W., & Zhang, Y. (2023). No (Cambrian) explosion and no (Ordovician) event: A single long-term radiation in the early Palaeozoic. Palaeogeography, Palaeoclimatology, Palaeoecology, 623, Article 111592. https://doi.org/10.1016/j.palaeo.2023.111592
• Harper, D. A., Meidla, T., & Servais, T. (2023). A short history of the Ordovician System: from overlapping unit stratotypes to global stratotype sections and points. Geological Society Special Publications, 532(1), https://doi.org/10.1144/sp532-2022-285
• Murray, J., MacGabhann, B. A., Doyle, E., Mángano, M. G., Tyrrell, S., & Harper, D. A. (2023). An enigmatic large discoidal fossil from the Pennsylvanian of County Clare, Ireland. Palaeoworld, https://doi.org/10.1016/j.palwor.2023.01.008
• Harper, D. A. (2023). The Irish Ordovician brachiopod fauna: A taxonomic renaissance. Estonian Journal of Earth Sciences, 72(1), 38-41. https://doi.org/10.3176/earth.2023.35
• Saleh, F., Vaucher, R., Vidal, M., Hariri, K. E., Laibl, L., Daley, A. C., …Lefebvre, B. (2022). New fossil assemblages from the Early Ordovician Fezouata Biota. Scientific Reports, 12(1), Article 20773. https://doi.org/10.1038/s41598-022-25000-z
• Aubry, M., Piller, W. E., Gibbard, P. L., Harper, D. A., & Finney, S. C. (2022). Ratification of subseries/subepochs as formal rank/units in international chronostratigraphy. Episodes, 45(1), 97-99. https://doi.org/10.18814/epiiugs/2021/021016
• Hughes, N. C., Peng, S., Harper, D. A., Myrow, P. M., Phạm, N. K., Wernette, S. J., & Zhu, X. (2022). Cambrian and earliest Ordovician fauna and geology of the Sông Đà and adjacent terranes in Việt Nam (Vietnam). Geological Magazine, 159(1), 55-80. https://doi.org/10.1017/s0016756821000844
• Pereira, S., Colmenar, J., Mortier, J., Vanmeirhaeghe, J., Verniers, J., Štorch, P., …Gutiérrez-Marco, J. C. (2021). Hirnantia Fauna from the Condroz Inlier, Belgium: another case of a relict Ordovician shelly fauna in the Silurian?. Journal of Paleontology, 6, 1189-1215. https://doi.org/10.1017/jpa.2021.74
• Zhang, M., Chen, Z., & Harper, D. A. (2021). An atypical Burgess Shale-type fossil assemblage from Cambrian Stage 4 of the Jingshan area, South China: Taphonomy, palaeoecology, and global correlations. Global and Planetary Change, 206, Article 103640. https://doi.org/10.1016/j.gloplacha.2021.103640
• Simonet Roda, M., Griesshaber, E., Angiolini, L., Harper, D. A., Jansen, U., Bitner, M. A., …Schmahl, W. W. (2021). The evolution of thecideide microstructures and textures: traced from Triassic to Holocene. Lethaia: An International Journal of Palaeontology and Stratigraphy, 54(4), 558-577. https://doi.org/10.1111/let.12422
• Harper, D. A., Stouge, S., Christiansen, J. L., Topper, T. P., Alwmark, C., Richoz, S., & Ahlberg, P. (2021). Early Cambrian brachiopod-dominated shell concentrations from North-East Greenland: Environmental and taphonomic implications. Global and Planetary Change, 204, Article 103560. https://doi.org/10.1016/j.gloplacha.2021.103560
• Donovan, S. K., Jones, S. J., King, A. R., & Harper, D. A. (2021). Pliocene trace fossils from oyster substrates in the Nijar Basin, Betic Cordillera, southern Spain. Proceedings of the Geologists' Association, 132(3), 358-368. https://doi.org/10.1016/j.pgeola.2021.02.004
• Guo, Z., Chen, Z., Harper, D. A., & Huang, Y. (2021). Permian–Triassic phylogenetic and morphologic evolution of rhynchonellide brachiopods. Paleobiology, https://doi.org/10.1017/pab.2021.25
• Servais, T., Cascales-Minana, B., & Harper, D. (2021). The Great Ordovician Biodiversification Event (GOBE) is not a single event. Paleontological Research, 25(4), 315-328. https://doi.org/10.2517/2021pr001
• Harper, D. A., Cascales-Miñana, B., Kroeck, D. M., & Servais, T. (2021). The palaeogeographical impact on the biodiversity of marine faunas during the Ordovician radiations. Global and Planetary Change, 207, Article 103665. https://doi.org/10.1016/j.gloplacha.2021.103665
• Jiayu, R., Harper, D., Bing, H., Rongyu, L., Xiaole, Z., & Di, C. (2020). The latest Ordovician Hirnantian brachiopod faunas: New global insights. Earth-Science Reviews, 208, Article 103280. https://doi.org/10.1016/j.earscirev.2020.103280
• Zhen, Y. Y., Zhang, Y., Harper, D. A., Zhan, R., Fang, X., Wang, Z., …Li, W. (2020). Ordovician successions in southern-central Xizang (Tibet), China—Refining the stratigraphy of the Himalayan and Lhasa terranes. Gondwana Research, 83, 372-389. https://doi.org/10.1016/j.gr.2020.01.023
• Rong, J., Aung, K. P., Zhan, R., Huang, B., Harper, D. A., Chen, D., …Zhang, X. (2020). The latest Ordovician Hirnantia brachiopod fauna of Myanmar: Significance of new data from the Mandalay Region. Palaeoworld, 29(1), 1-30. https://doi.org/10.1016/j.palwor.2019.07.002
• Huang, B., Rong, J., Harper, D. A., & Zhou, H. (2020). A nearshore Hirnantian brachiopod fauna from South China and its ecological significance. Journal of Paleontology, 94(2), 239-254. https://doi.org/10.1017/jpa.2019.90
• Huang, B., Zhou, H., Harper, D. A., Zhan, R., Zhang, X., Chen, D., & Rong, J. (2020). A latest Ordovician Hirnantia brachiopod fauna from western Yunnan, Southwest China and its paleobiogeographic significance. Palaeoworld, 29(1), 31-46. https://doi.org/10.1016/j.palwor.2019.03.002
• Harper, D. A., Cascales-Miñana, B., & Servais, T. (2020). Early Palaeozoic diversifications and extinctions in the marine biosphere: a continuum of change. Geological Magazine, 157, 5-21. https://doi.org/10.1017/s0016756819001298
• Guo, Z., Chen, Z., & Harper, D. A. (2020). The Anisian (Middle Triassic) brachiopod fauna from Qingyan, Guizhou, south-western China. Journal of Systematic Palaeontology, 18(8), 647-701. https://doi.org/10.1080/14772019.2019.1682695
• curiosity. Irish Journal of Earth Sciences, 38, 73-90. https://doi.org/10.3318/ijes.2020.38.5
• Stouge, S., Harper, D. A., Zhan, R., Liu, J., & Stemmerik, L. (2020). Middle Ordovician (Darriwilian) conodonts from southern Tibet, the Indian passive margin: implications for the age and correlation of the roof of the world. Geological Magazine, 1-25. https://doi.org/10.1017/s0016756820001077
• Rønning, K. J., Bruton, D. L., Harper, D. A., Høyberget, M., Maletz, J., & Nakrem, H. A. (2020). A Cambrian–Ordovician boundary section in the Rafnes–Herøya submarine tunnel, Skien–Langesund District, southern Norway. Norsk Geologisk Tidsskrift, 100(1), 1-24. https://doi.org/10.17850/njg100-3-3
• Ling, M., Zhan, R., Wang, G., Wang, Y., Amelin, Y., Tang, P., …Rong, J. (2019). An extremely brief end Ordovician mass extinction linked to abrupt onset of glaciation. Solid earth sciences, 4(4), 190-198. https://doi.org/10.1016/j.sesci.2019.11.001
• Saleh, F., Candela, Y., Harper, D. A., Polechová, M., Lefebvre, B., & Pittet, B. (2019). Storm-induced community dynamics in the Fezouata Biota (Lower Ordovician, Morocco). PALAIOS, 33(12), 535-541. https://doi.org/10.2110/palo.2018.055
• Servais, T., Cascales-Miñana, B., Cleal, C. J., Gerrienne, P., Harper, D. A., & Neumann, M. (2019). Revisiting the Great Ordovician Diversification of land plants: Recent data and perspectives. Palaeogeography, Palaeoclimatology, Palaeoecology, 534, https://doi.org/10.1016/j.palaeo.2019.109280
• Harper, D., Hammarlund, E., Topper, T., Nielsen, A., Rasmussen, J., T-y.s., P., & Smith, M. (2019). The Sirius Passet Lagerstätte of North Greenland: a remote window on the Cambrian Explosion. Journal of the Geological Society, 176(6), 1023-1037. https://doi.org/10.1144/jgs2019-043
• Angiolini, L., Crippa, G., Azmy, K., Capitani, G., Confalonieri, G., Della Porta, G., …Stephenson, M. H. (2019). The giants of the phylum Brachiopoda: a matter of diet?. Palaeontology, 62(6), 889-917. https://doi.org/10.1111/pala.12433
• Schmitz, B., Farley, K. A., Goderis, S., Heck, P. R., Bergström, S. M., Boschi, S., …Terfelt, F. (2019). An extraterrestrial trigger for the mid-Ordovician ice age: Dust from the breakup of the L-chondrite parent body. Science Advances, 5(9), Article eaax4184. https://doi.org/10.1126/sciadv.aax4184
• Huang, B., Harper, D. A., Zhou, H., Zhan, R., Wang, Y., Tang, P., …Rong, J. (2019). A new Cathaysiorthis (Brachiopoda) fauna from the lower Llandovery of eastern Qinling, China. Papers in Palaeontology, 5(3), 537-557. https://doi.org/10.1002/spp2.1253
• Harper, D. (2019). The Golden Spike Still Glitters: The (Re)construction of a Global Chronostratigraphy. Acta Geologica Sinica - English Edition, 93(S3), 24-27. https://doi.org/10.1111/1755-6724.14234
• Harper, D. A., Topper, T. P., Cascales-Miñana, B., Servais, T., Zhang, Y., & Ahlberg, P. (2019). The Furongian (late Cambrian) Biodiversity Gap: Real or apparent?. Palaeoworld, 28(1-2), 4-12. https://doi.org/10.1016/j.palwor.2019.01.007
• Hammarlund, E. U., Smith, M. P., Rasmussen, J. A., Nielsen, A. T., Canfield, D. E., & Harper, D. A. (2019). The Sirius Passet Lagerstätte of North Greenland—A geochemical window on early Cambrian low‐oxygen environments and ecosystems. Geobiology, 17(1), 12-26. https://doi.org/10.1111/gbi.12315
• Huang, B., Harper, D. A., Zhou, H., & Rong, J. (2019). From shallow to deep water: an ecological study of the Hirnantia brachiopod Fauna (Late Ordovician) and its global implications. Lethaia: An International Journal of Palaeontology and Stratigraphy, 1-13. https://doi.org/10.1111/let.12360
• Drost, K., Chew, D., Petrus, J. A., Scholze, F., Woodhead, J. D., Schneider, J. W., & Harper, D. A. (2018). An Image Mapping Approach to U-Pb LA-ICP-MS Carbonate Dating and Applications to Direct Dating of Carbonate Sedimentation. Geochemistry, Geophysics, Geosystems, 19(12), 4631-4648. https://doi.org/10.1029/2018gc007850
• Bogolepova, O. K., Donovan, S. K., Harper, D. A., Suyarkova, A. A., Yakupov, R., & Gubanov, A. P. (2018). New records of brachiopods and crinoids from the Silurian (Wenlock) of the southern Urals, Russia. GFF, 140(4), 323-331. https://doi.org/10.1080/11035897.2018.1526210
• Hints, L., Harper, D., & Paškevičius, J. (2018). Diversity and biostratigraphic utility of Ordovician brachiopods in the East Baltic. Estonian Journal of Earth Sciences, 67(3), 176-191. https://doi.org/10.3176/earth.2018.14
• Strang, K., Harper, D., Donovan, S., & Portell, R. (2018). Silicification of low-magnesium mollusc shells from the Upper Oligocene of Antigua, Lesser Antilles
• Servais, T., & Harper, D. A. (2018). The Great Ordovician Biodiversification Event (GOBE): definition, concept and duration. Lethaia: An International Journal of Palaeontology and Stratigraphy, 51(2), 151-164. https://doi.org/10.1111/let.12259
• Pohl, A., Harper, D. A., Donnadieu, Y., Le Hir, G., Nardin, E., & Servais, T. (2018). Possible patterns of marine primary productivity during the Great Ordovician Biodiversification Event. Lethaia: An International Journal of Palaeontology and Stratigraphy, 51(2), 187-197. https://doi.org/10.1111/let.12247
• Topper, T. P., Greco, F., Hofmann, A., Beeby, A., & Harper, D. A. (2018). Characterization of kerogenous films and taphonomic modes of the Sirius Passet Lagerstätte, Greenland. Geology, 46(4), 359-362. https://doi.org/10.1130/g39930.1
• Harper, D., & Servais, T. (2018). Contextualizing the Onset of the Great Ordovician Biodiversification Event. Lethaia: An International Journal of Palaeontology and Stratigraphy, 51(2), 149-150. https://doi.org/10.1111/let.12264
• Topper, T., Zhang, Z., Gutiérrez-Marco, J., & Harper, D. (2018). The dawn of a dynasty: life strategies of Cambrian and Ordovician brachiopods. Lethaia: An International Journal of Palaeontology and Stratigraphy, 51(2), 254-266. https://doi.org/10.1111/let.12229
• Park, T. S., Kihm, J., Woo, J., Park, C., Lee, W. Y., Smith, M. P., …Vinther, J. (2018). Brain and eyes of Kerygmachela reveal protocerebral ancestry of the panarthropod head. Nature Communications, 9(1), Article 1019. https://doi.org/10.1038/s41467-018-03464-w
• Erlykin, A. D., Harper, D. A., Sloan, T., & Wolfendale, A. W. (2018). Periodicity in extinction rates. Palaeontology, 61(1), 149-158. https://doi.org/10.1111/pala.12334
• Blake, D. B., Donovan, S. K., & Harper, D. A. (2017). A new Silurian ophiuroid from the west of Ireland. Irish Journal of Earth Sciences, 35, 57-66. https://doi.org/10.3318/ijes.2017.35.57
• Finnegan, S., Rasmussen, C., & Harper, D. (2017). Identifying the most surprising victims of mass extinction events: an example using Late Ordovician Brachiopods. Biology Letters, 13(9), Article 2017400. https://doi.org/10.1098/rsbl.2017.0400
• Nielsen, M., Rasmussen, J., & Harper, D. (2017). Sexual dimorphism within the stem-group arthropod Isoxys volucris from the Sirius Passet Lagerstätte, North Greenland. Bulletin of the Geological Society of Denmark, 65, 47-58
• Harper, D. A., Popov, L. E., & Holmer, L. E. (2017). Brachiopods: origin and early history. Palaeontology, 60(5), 609-631. https://doi.org/10.1111/pala.12307
• Harper, D. A., Parkes, M. A., & Ren-Bin, Z. (2017). Late Ordovician deep-water brachiopod fauna from Raheen, Waterford Harbour, Ireland. Irish Journal of Earth Sciences, 35, 1-18. https://doi.org/10.3318/ijes.2017.35.1
• Donovan, S. K., Harper, D. A., Portell, R. W., & Toomey, J. K. (2017). Echinoids as hard substrates: varied examples from the Oligocene of Antigua, Lesser Antilles. Proceedings of the Geologists' Association, 128(3), 326-331. https://doi.org/10.1016/j.pgeola.2017.04.003
• Huang, B., Harper, D. A., Rong, J., & Zhan, R. (2017). Brachiopod faunas after the end Ordovician mass extinction from South China: Testing ecological change through a major taxonomic crisis. Journal of Asian Earth Sciences, 138, 502-514. https://doi.org/10.1016/j.jseaes.2017.02.043
• Erlykin, A. D., Harper, D. A., Sloan, T., & Wolfendale, A. W. (2017). Mass extinctions over the last 500 myr: an astronomical cause?. Palaeontology, 60(2), 159-167. https://doi.org/10.1111/pala.12283
• Donovan, S., Harper, D., & Portell, R. (2017). Shell-Filled Burrows in the Upper Oligocene Antigua Formation, Antigua, Lesser Antilles. Ichnos: an International Journal of Plant and Animal Traces, 24, 72-77. https://doi.org/10.1080/10420940.2016.1223653
• Strang, K., Armstrong, H., & Harper, D. (2016). Minerals in the gut: scoping a Cambrian digestive system. Royal Society Open Science, 3(11), Article 160420. https://doi.org/10.1098/rsos.160420
• Donovan, S., Jones, B., & Harper, D. (2016). Neogene echinoids from the Cayman Islands, West Indies: regional implications. Geological Journal, 51(6), 864-879. https://doi.org/10.1002/gj.2703
• Strang, K., Armstrong, H., Harper, D., & Trabucho-Alexandre, J. P. (2016). The Sirius Passet Lagerstatte: silica death masking opens the window on the earliest matground community of the Cambrian explosion. Lethaia: An International Journal of Palaeontology and Stratigraphy, 49(4), 631-643. https://doi.org/10.1111/let.12174
• Chenyi, T., Zhong-Qiang, C., & Harper, D. (2016). Permian–Triassic evolution of the Bivalvia: Extinction-recovery patterns linked to ecologic and taxonomic selectivity. Palaeogeography, Palaeoclimatology, Palaeoecology, 459, 53-62. https://doi.org/10.1016/j.palaeo.2016.06.042
• Jensen, S., Harper, D., & Stouge, S. (2016). Trace fossils from the lower Cambrian Kløftelv Formation, Ella Ø, North-East Greenland. GFF, 138(3), 369-376. https://doi.org/10.1080/11035897.2015.1076029
• their palaeogeographic implication. Geological Journal, 51(4), 584-599. https://doi.org/10.1002/gj.2659
• Jakobsen, K., Brock, G., Nielsen, A., & Harper, D. (2016). A Darriwilian (Middle Ordovician) bivalve-dominated molluscan fauna from the Stairway Sandstone, Amadeus Basin, central Australia. Acta Palaeontologica Polonica, 61(4), 897-924. https://doi.org/10.4202/app.00215.2015
• Finnegan, S., Rasmussen, C., & Harper, D. (2016). Biogeographic and bathymetric determinants of brachiopod extinction and survival during the Late Ordovician mass extinction. Proceedings of the Royal Society B: Biological Sciences, 283(1829), Article 20160007. https://doi.org/10.1098/rspb.2016.0007
• Harper, D., & Hints, L. (2016). Hirnantian (Late Ordovician) brachiopod faunas across Baltoscandia: A global and regional context. Palaeogeography, Palaeoclimatology, Palaeoecology, 444, 71-83. https://doi.org/10.1016/j.palaeo.2015.11.044
• Rasmussen, C. Ø., Ullmann, C., Jakobsen, K., Lindskog, A., Hansen, J., Hansen, T., …Harper, D. (2016). Onset of main Phanerozoic marine radiation sparked by emerging Mid Ordovician icehouse. Scientific Reports, 6, Article 18884. https://doi.org/10.1038/srep18884
• Rohr, D., Harper, D., Stouge, S., & Christiansen, J. (2015). Ordovician Gastropoda from Northeast Greenland. Bulletin of Geosciences (On-line), 90, 795-805. https://doi.org/10.3140/bull.geosci.1566
• Donovan, S., Harper, D., & Portell, R. (2015). In deep water: a crinoid–brachiopod association in the Upper Oligocene of Antigua, West Indies. Lethaia: An International Journal of Palaeontology and Stratigraphy, 48(3), 291-298. https://doi.org/10.1111/let.12105
• Zhan, R., Jin, J., & Harper, D. (2015). Ecosystem revolution and evolution in the Early–Mid Paleozoic. Palaeoworld, 24(1-2), 1-4. https://doi.org/10.1016/j.palwor.2015.05.001
• the Upper Ordovician of North America. Papers in Palaeontology, 1, 237-253. https://doi.org/10.1002/spp2.1014
• Jakobsen, K., Brock, G., Nielsen, A., Mathieson, D., & Harper, D. (2015). Brachiopods associated with stromatoporoid mounds from the Middle to Upper Ordovician Cashions Creek Limestone, Tasmania. Memoir ... of the Association of Australasian Palaeontologists, 45, 249-265
• Hints, L., & Harper, D. (2015). The Hirnantian (Late Ordovician) brachiopod fauna of the East Baltic: Taxonomy of the key species. Acta Palaeontologica Polonica, 60(2), 395-420. https://doi.org/10.4202/app.2013.0010
• Harper, D., Hammarlund, E., & Rasmussen, C. (2014). End Ordovician extinctions : a coincidence of causes. Gondwana Research, 25(4), 1294-1307. https://doi.org/10.1016/j.gr.2012.12.021
• Jagt, J., Thuy, B., Donovan, S., Stohr, S., Portell, R., Pickerill, R., …Jackson, T. (2014). A starfish bed in the Middle Miocene Grand Bay Formation of Carriacou, The Grenadines (West Indies). Geological Magazine, 151(3), 381-393. https://doi.org/10.1017/s0016756813000204
• Vinther, J., Stein, M., Longrich, N., & Harper, D. (2014). A suspension-feeding anomalocarid from the Early Cambrian. Nature, 507(7493), 496-499. https://doi.org/10.1038/nature13010
• Colmenar, J., Harper, D., & Villas, E. (2014). Morphofunctional analysis of Svobodaina species (Brachiopoda, Heterorthidae) from South-Western Europe. Palaeontology, 57(1), 193-214. https://doi.org/10.1111/pala.12061
• Bing, H., Harper, D., & Renbin, Z. (2014). Test of sampling sufficiency in palaeontology. GFF, 136(1), 105-109. https://doi.org/10.1080/11035897.2014.882976
• Harper, D., Jisuo, J., & Rasmussen, C. (2014). Late Ordovician carbonate mounds from North Greenland: a peri-Laurentian dimension to the Boda Event?. GFF, 136(1), 95-99. https://doi.org/10.1080/11035897.2013.865669
• Servais, T., Danielian, T., Harper, D., & Munnecke, A. (2014). Possible oceanic circulation patterns, surface water currents and upwelling zones in the Early Palaeozoic. GFF, 136(1), 229-233. https://doi.org/10.1080/11035897.2013.876659
• Renbin, Z., Harper, D., Jisuo, J., Yan, L., Jianbo, L., Stemmerik, L., & Stouge, S. (2014). Middle Ordovician Aporthophyla brachiopod fauna from the roof of the World, Southern Tibet. Palaeontology, 57(1), 141-170. https://doi.org/10.1111/pala.12058
• Jakobsen, K., Nielsen, A., Harper, D., & Brock, G. (2014). Trilobites from the Middle Ordovician Stairway Sandstone, Amadeus Basin, central Australia. Alcheringa: An Australasian Journal of Palaeontology, 38, 70-96. https://doi.org/10.1080/03115518.2013.828253
• Oligocene oysters, Antigua, West Indies. Proceedings of the Geologists' Association, 125(1), 99-106. https://doi.org/10.1016/j.pgeola.2013.10.002
• Smith, M., & Harper, D. (2013). Causes of the Cambrian Explosion. Science, 341(6152), 1355-1356. https://doi.org/10.1126/science.1239450
• Stein, M., Budd, G., Peel, J., & Harper, D. (2013). Arthroaspis n. gen., a common element of the Sirius Passet Lagerstätte (Cambrian, North Greenland), sheds light on trilobite ancestry. BMC Evolutionary Biology, 13, Article 99. https://doi.org/10.1186/1471-2148-13-99
• Servais, T., Cecca, F., Harper, D., Isozaki, Y., & Mac Niocaill, C. (2013). Palaeozoic palaeogeographical and palaeobiogeographical nomenclature. Memoirs, 38(1), 25-33. https://doi.org/10.1144/m38.3
• Zuykov, M., Pelletier, E., & Harper, D. (2013). Bivalve mollusks in metal pollution studies: from bioaccumulation to biomonitoring. Chemosphere, 93, 201-208. https://doi.org/10.1016/j.chemosphere.2013.05.001
• Topper, T., Harper, D., & Brock, G. (2013). Ancestral billingsellides and the evolution and phylogenetic relationships of early rhynchonelliform brachiopods. Journal of Systematic Palaeontology, 11(7), 821-833. https://doi.org/10.1080/14772019.2012.728253
• Jakobsen, K., Brock, G., Nielsen, A., Topper, T., & Harper, D. (2013). Middle Ordovician brachiopods from the Stairway Sandstone, Amadeus Basin, central Australia. Alcheringa: An Australasian Journal of Palaeontology, 38, 190-208. https://doi.org/10.1080/03115518.2013.858204
• rhynchonelliform brachiopods. Memoirs, 38(1), 127-144. https://doi.org/10.1144/m38.11
• Tvären crater system, Sweden. Palaeogeography, Palaeoclimatology, Palaeoecology, 369, 114-124. https://doi.org/10.1016/j.palaeo.2012.10.008
• Jisuo, J., Harper, D., Cocks, L., McCausland, P., Rasmussen, C., & Sheehan, P. (2013). Precisely locating the Ordovician equator in Laurentia. Geology, 41(2), 107-110. https://doi.org/10.1130/g33688.1
• 150 years of confusion. GFF, 135(2), 191-203. https://doi.org/10.1080/11035897.2013.811440
• Harper, D., & Servais, T. (2013). Early Palaeozoic biogeography and palaeogeography: towards a modern synthesis. Memoirs, 38(1), 1-3. https://doi.org/10.1144/m38.1
• Topper, T., Skovsted, C., Harper, D., & Ahlberg, P. (2013). A Bradoriid and Brachiopod Dominated Shelly Fauna from the Furongian(Cambrian) of Västergötland, Sweden. Journal of Paleontology, 87(1), 69-83. https://doi.org/10.1666/12-047r.1
• Topper, T., Skovsted, C., Peel, J., & Harper, D. (2013). Moulting in the lobopodian Onychodictyon from the lower Cambrian of Greenland. Lethaia: An International Journal of Palaeontology and Stratigraphy, 46, 490-495. https://doi.org/10.1111/let.12026
• Bing, H., Harper, D., & Hammer, O. (2013). Introduction to PAST, a comprehensive statistics software package for paleontological data analysis. Gushengwu xuebao, 52, 161-181
• Mangano, M., Bromley, R., Harper, D., Nielsen, A., Smith, M., & Vinther, J. (2012). Nonbiomineralized carapaces in Cambrian seafloor landscapes (Sirius Passet, Greenland): Opening a new window into early Phanerozoic benthic ecology. Geology, 40(6), 519-522. https://doi.org/10.1130/g32853.1
• Hammarlund, E., Dahl, T., Harper, D., Bond, D., Nielsen, A., Bjerrum, C., …Canfield, D. (2012). A sulfidic driver for the end-Ordovician mass extinction. Earth and Planetary Science Letters, 331-332, 128-139. https://doi.org/10.1016/j.epsl.2012.02.024
• Donovan, S., Harper, D., Renbin, Z., Stemmerik, L., Jianbo, L., & Stouge, S. (2012). A primitive cladid crinoid from the Jiacun Group, Tibet (Darriwilian, Middle Ordovician). Geological Journal, 47, 653-660. https://doi.org/10.1002/gj.2436
• of Laurentia. Palaeogeography, Palaeoclimatology, Palaeoecology, 367-368, 73-88. https://doi.org/10.1016/j.palaeo.2011.05.023
• Zuykov, M., Vinarski, M., Pelletier, E., Demers, S., & Harper, D. (2012). Shell malformations in seven species of pond snail (Gastropoda, Lymnaeidae): analysis of large museum collections. Zoosystematics and Evolution, 88(2), 365-368. https://doi.org/10.1002/zoos.201200025
• Topper, T., Holmer, L., Skovsted, C., Brock, G., Balthasar, U., Larsson, C., …Harper, D. (2012). The oldest brachiopods from the lower Cambrian of South Australia. Acta Palaeontologica Polonica, 58(1), 93-109. https://doi.org/10.4202/app.2011.0146
• Zuykov, M., Harper, D., & Pelletier, E. (2011). Revision of the Ordovician brachiopod genus Noetlingia Hall and Clarke, 1893. Journal of Paleontology, 85(3), 595-598
• Rasmussen, C., & Harper, D. (2011). Did the amalgamation of continents drive the end Ordovician mass extinctions?. Palaeogeography, Palaeoclimatology, Palaeoecology, 311(1-2), 48-62. https://doi.org/10.1016/j.palaeo.2011.07.029
• Rasmussen, C., & Harper, D. (2011). Interrogation of distributional data for the End Ordovician crisis interval: where did disaster strike?. Geological Journal, 46, 478-500. https://doi.org/10.1002/gj.1310
• Vinther, J., Smith, M., & Harper, D. (2011). Vetulicolians from the Lower Cambrian Sirius Passet Lagerstatte, North Greenland, and the polarity of morphological characters in basal deuterostomes. Palaeontology, 54(3), 711-719. https://doi.org/10.1111/j.1475-4983.2011.01034.x
• Frisk, Å., & Harper, D. (2011). Palaeoenvironmental aspects of Late Ordovician Sericoidea shell concentrations in an impact crater, Tvaren, Sweden. Lethaia: An International Journal of Palaeontology and Stratigraphy, 44(4), 383-396. https://doi.org/10.1111/j.1502-3931.2010.00250.x
• Vinther, J., Eibye-Jacobsen, D., & Harper, D. (2011). An Early Cambrian stem polychaete with pygidial cirri. Biology Letters, 7(6), 929-932. https://doi.org/10.1098/rsbl.2011.0592
• Stouge, S., Christiansen, J., Harper, D., Houmark-Nielsen, M., Kristiansen, K., MacNiocaill, C., & Buchard-Westergaard, B. (2011). Neoproterozoic (Cryogenian–Ediacaran) deposits in East and North-East Greenland. Memoirs, 36(1), 581-592. https://doi.org/10.1144/m36.56
• of the Chernobyl NPP (Ukraine)?. Environmentalist, 31(4), 369-375. https://doi.org/10.1007/s10669-011-9347-4
• Hughes, N., Myrow, P., McKenzie, N., Harper, D., Bhargava, O., Tangri, S., …Fanning, C. (2011). Cambrian rocks and faunas of the Wachi La, Black Mountains, Bhutan. Geological Magazine, 148(03), 351-379. https://doi.org/10.1017/s0016756810000750
• Munnecke, A., Calner, M., & Harper, D. (2010). How does sea level correlate with sea-water chemistry? A progress report from the Ordovician and Silurian. Palaeogeography, Palaeoclimatology, Palaeoecology, 296(3-4), 213-216. https://doi.org/10.1016/j.palaeo.2010.07.009
• Azmy, K., Stouge, S., Christiansen, J., Harper, D., Knight, I., & Boyce, D. (2010). Carbon-isotope stratigraphy of the Lower Ordovician succession in Northeast Greenland: Implications for correlations with St. George Group in western Newfoundland (Canada) and beyond. Sedimentary Geology, 225(1-2), 67-81. https://doi.org/10.1016/j.sedgeo.2010.01.007
• Munnecke, A., Calner, M., Harper, D., & Servais, T. (2010). Ordovician and Silurian sea-water chemistry, sea level, and climate: A synopsis. Palaeogeography, Palaeoclimatology, Palaeoecology, 296(3-4), 389-413. https://doi.org/10.1016/j.palaeo.2010.08.001
• Rasmussen, C., Ebbestad, J., & Harper, D. (2010). Unravelling a Late Ordovician pentameride (Brachiopoda) hotspot from the Boda Limestone, Siljan district, central Sweden. GFF, 132(3-4), 133-152. https://doi.org/10.1080/11035897.2010.506008
• Candela, Y., & Harper, D. (2010). Late Ordovician (Katian) brachiopods from the Southern Uplands of Scotland: biogeographic patterns on the edge of Laurentia. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 100(03), 253-274. https://doi.org/10.1017/s1755691010008121
• Bing, H., Harper, D., Renbin, Z., & Jiayu, R. (2010). Can the Lilliput Effect be detected in the brachiopod faunas of South China following the terminal Ordovician mass extinction?. Palaeogeography, Palaeoclimatology, Palaeoecology, 285(3-4), 277-286. https://doi.org/10.1016/j.palaeo.2009.11.020
• Harper, D., Parkes, M., & McConnell, B. (2010). Late Ordovician (Sandbian) brachiopods from the Mweelrea Formation, South Mayo, western Ireland: stratigraphic and tectonic implications. Geological Journal, 45, 445-450. https://doi.org/10.1002/gj.1210
• Servais, T., Owen, A., Harper, D., Kroger, B., & Munnecke, A. (2010). The Great Ordovician Biodiversification Event (GOBE): The palaeoecological dimension. Palaeogeography, Palaeoclimatology, Palaeoecology, 294(3-4), 99-119. https://doi.org/10.1016/j.palaeo.2010.05.031
• Dahlqvist, P., Harper, D., & Wickstrom, L. (2010). Late Ordovician shelly faunas from Jamtland: palaeocommunity development along the margin of the Swedish Caledonides. Bulletin of Geosciences (On-line), 85, 505-512. https://doi.org/10.3140/bull.geosci.1175
• Jin, J., Harper, D., & Rasmussen, C. (2009). Sulcipentamerus (Pentamerida, Brachiopoda) from the Lower Silurian Washington Land Group, North Greenland. Palaeontology, 52(2), 385-399. https://doi.org/10.1111/j.1475-4983.2009.00849.x
• Rasmussen, C., Nielsen, A., & Harper, D. (2009). Ecostratigraphical interpretation of lower Middle Ordovician East Baltic sections based on brachiopods. Geological Magazine, 146(05), 717-731. https://doi.org/10.1017/s0016756809990148
• Schmitz, B., Harper, D., Peucker-Ehrenbrink, B., Stouge, S., Alwmark, C., Cronholm, A., …Xiaofeng, W. (2008). Asteroid breakup linked to the Great Ordovician Biodiversification Event. Nature Geoscience, 1(1), 49-53. https://doi.org/10.1038/ngeo.2007.37
• Harper, D., & Pickerill, R. (2008). Generation of brachiopod-dominated shell beds in the Miocene rocks of Carriacou, Lesser Antilles. Geological Journal, 43(5), 573-581. https://doi.org/10.1002/gj.1131